5.2.8 · Biology › Population & Community Ecology
Competition ek biological struggle hai jo tab hoti hai jab do ya zyada organisms ko ek hi limited resource ki zaroorat hoti hai, ek hi time par, ek hi jagah par. Isse ek "minus-minus" interaction (−/−) samjho: dono competitors lose karte hain, agar compare karo us situation se jahan resources unlimited hote. Key insight yeh hai: resources finite hote hain, demand supply se zyada hoti hai, aur evolution inefficiency ko punish karta hai.
Competition kyun matter karti hai: Yeh community structure ko shape karne wali sabse powerful forces mein se ek hai — natural selection drive karti hai, population growth ko limit karti hai, aur yeh decide karti hai ki kaun si species coexist kar sakti hain.
Organisms ke beech ek aisa interaction (same ya alag species ka) jisme dono ki fitness reduce ho jaati hai (survival, growth, ya reproduction) kyunki dono ko ek limited supply wale resource ki zaroorat hoti hai.
Resource ho sakta hai:
Consumable : khaana, paani, nutrients, prey (ek baar use hone ke baad, khatam)
Non-consumable : jagah, nesting sites, mates, sunlight (monopolize ho sakta hai, lekin "khaya" nahi ja sakta)
Mechanism: Jab individual A resource R ki ek unit use karta hai, toh individual B ke liye R kam ho jaata hai. Isse ek negative feedback loop banta hai jo population growth ko slow karta hai jab density badhti hai.
==Intraspecific competition == = competition within ek species ke andar (same species ke members compete karte hain)
==Interspecific competition == = competition between alag-alag species ke beech
Intuition Intraspecific Usually Zyada Strong Kyun Hoti Hai
Same species ke members ke identical niche requirements hote hain — woh same khaana khaate hain, same breeding sites chahiye, same microhabitat mein rehte hain. Woh perfect competitors hain. Isliye density-dependent regulation kaam karti hai: jab tumhari apni population badhti hai, tum sabse zyada apni hi tarah ke logon se compete karte ho .
Exponential growth model se shuru karo (unlimited resources):
d t d N = r N
jahan r = per capita growth rate (constant), N = population size.
Problem: Yeh infinite growth predict karta hai. Reality? Jab N badhta hai, resources per individual kam hote hain, toh r ko decline karna padta hai.
Step 1: r ko density-dependent banao. Sabse simple linear form:
r a c t u a l = r 0 ( 1 − K N )
jahan:
r 0 = intrinsic growth rate (maximum, jab N ≈ 0 )
K = carrying capacity (maximum N jo environment sustain kar sakta hai)
K N = carrying capacity ka kitna proportion use ho chuka hai
Yeh form kyun? Jab N = 0 , koi competition nahi, toh r = r 0 . Jab N = K , toh r = 0 (growth rukti hai). Jab N > K , toh r < 0 (population decline karti hai).
Step 2: Growth equation mein substitute karo:
d t d N = r 0 N ( 1 − K N )
Yeh ==logistic growth equation == hai — intraspecific competition ka mathematical signature.
Worked example Worked Example 1: Yeast in a Flask
Ek lab culture of yeast N 0 = 10 cells se shuru hoti hai. Carrying capacity K = 1000 cells hai, aur r 0 = 0.5 hr − 1 hai.
Calculate karo: Growth rate jab N = 100 , = 500 , aur N = 900 .
Solution:
N = 100 par:
d t d N = 0.5 × 100 × ( 1 − 1000 100 ) = 50 × 0.9 = 45 cells/hr
Yeh step kyun? Jab population chhoti hai (K ka 10%), intraspecific competition weak hai — 90% resources abhi bhi available hain.
N = 500 par:
d t d N = 0.5 × 500 × ( 1 − 1000 500 ) = 250 × 0.5 = 125 cells/hr
N = 100 se zyada fast kyun? Competition strong ho gayi hai (sirf 50% resources bache hain), lekin badi population base (N = 500 , 100 ki jagah) isse compensate kar leti hai. Yeh peak growth rate hai.
N = 900 par:
d t d N = 0.5 × 900 × ( 1 − 1000 900 ) = 450 × 0.1 = 45 cells/hr
Yeh step kyun? Competition intense hai (K ka 90% use ho chuka hai). Growth dramatically slow ho jaati hai bade N ke bawajood.
==Exploitation competition == (scramble): Sabhi individuals ko resources tak equal access hota hai; woh bas consume karte hain. Sabko kam milta hai. Example: grass plants jo soil nitrogen ke liye compete karti hain — saari roots absorb karti hain, kissi ko poora nahi milta.
==Interference competition == (contest): Direct antagonistic interactions. Dominants resources monopolize karte hain; subordinates ko bahut kam ya kuch nahi milta. Example: territorial birds — winners breed karte hain, losers nahi. Hierarchies banti hain.
Worked example Worked Example 2: Tadpole Density Experiment
Ek ecologist tadpoles ko tanks mein 5, 20, 50, 100 per liter densities par rakhti hai. Woh metamorphosis par average mass measure karti hai:
Density (tadpoles/L)
Avg mass (g)
Survival (%)
5
2.4
95
20
1.8
88
50
1.1
70
100
0.6
45
Interpretation:
Mass decline = algae (khaane) ke liye exploitation competition. Density badhne par har tadpole ko kam milta hai.
Survival decline = jab competition extreme hoti hai, kuch tadpole starve kar jaate hain.
Yeh kyun matter karta hai: Kam mass → chhote frogs → kam reproductive success. Intraspecific competition population ko reduced fecundity aur increased mortality ke zariye regulate karti hai.